SYSTEMATIC BIOLOGY

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SUPERCLASS CONODONTOMORPHI+

Conodontomorphi (KON-o-don-to-MOPF-e) is formed from three Greek roots meaning "cone-toothed forms" [cone- konos (κώνος); tooth- donti (δόντι); form- morfi (μορφή)].  The name is derived from denticles that are very abundant in Paleozoic strata.

The conodonts were very similar to lampreys in overall body form.  They had two large eyes and in-line fins only.  Their jawless mouths had batteries of teeth that seemed to serve the functions of grasping and then chopping prey.  Only in a few sites have the carbonized bodies of conodonts been found; so, the taxonomic system of the animals rests entirely on the structure of the teeth.  Their biology is a little difficult to infer from the scant evidence that we have today.  They were free-swimming and likely were predators (or at least scavengers) much like the hagfishes.  Their particular position in the Craniata is still in doubt.  Donoghue et al. (2000) placed them in the Subphylum Conodontophorida as distinct from the Vertebrata (craniata) and sisters to the Myxiniodea. 

HIERARCHICAL TAXONOMY OF THE SUPERCLASS CONODONTOMORPHI. The following descriptions come from Benton (2005) and Nelson (2006).  The structure of the following system is based on Sweet and Donoghue (2001).  Taxa in red are extinct.

Conodont teeth from http://energy.usgs.gov/images/oilgas/ha000001_sm.jpg

SUPERCLASS CONODONTOMORPHI+

CLASS CONODONTA                                 

Conodont teeth are common from the late Cambrian to the Late Triassic.  Though common, indeed so common that they were used as stratographic standards, the nature of the conodont organism was in doubt until 1983 (Briggs et al. 1983) when lamprey-like animals with conodont teeth were found as nearly complete and highly detailed carbonized films in Scotland.  They seem to have had paired eyes.  Their dental elements came in 3 forms called S, M, and P.  S and M elements were associated with the front end of the mouth and served to grasp prey.  P elements were at the back of the pharynx and served to chop the prey to pieces.  Although many genera are recognized, they are based on tooth morphology; so, relationships between the taxa are artificial.

Late Cambrian to late Triassic

UNASSIGNED GENERA

Acodus, Amorphognathus, Anticostiodus, Basselodus, Belodina, Carniodus, Cornuodus, Culumbodina, Curtognathus, Dapsilodus, Decoriconus, Diaphorodus, Drepanodus, Erismodus, Erraticodon, Fahraeusodus, Gamachignathus, Hamarodus, Histiodella, Istorinus, Juanognathus, Jumudontus, Kallidontus, Lenodus, Microzarkodina, Oepikodus, Oistodus, Oulodus, Paroistodus, Pedavis, Phragmodus, Polonodus, Pseudobelodina, Pseudooneotodus, Pteracontiodus, Pterospathodus, Pygodus, Scabbardella, Scolopodus, Scyphoidus, Spinodus, Staufferella, Stereoconus, Strachanognathus, Streptognathus, Taoqupognathus, Tripodus, Walliserodus, Yaoxianognathus.

? Semiaccontiodus, Clavohamulus, Drepanoistidus, Utahconus,  Hirsutodontus, Clydagnathus

UNASSIGNED FAMILY

FAMILY PARACORDYLODONTIDAE

Paracordylodus.

SUBCLASS CAVIDONTI

ORDER PROCONODONTIDA

UNASSIGNED GENERA

Proconodontus

FAMILY FRYXELLODONTIDAE

Dzikodus, Kallidontus.

FAMILY PYGODONTIDAE

FAMILY CORDYLODONTIDAE

Cordylodus, Econodontus, Iapetognathus.

FAMILY ASELLIIDAE

FAMILY DAPSILODONTIDAE

FAMILY BELODELLIDAE

SUBCLASS CONODONTI

ORDER PROTOPANDERODONTIDA

FAMILY PROTOPANDERODONTIDAE

Protopanderodus.

FAMILY CLAVOHAMULIDAE

FAMILY DREPANOISTODONTIDAE

Drepanoistododus

FAMILY ACANTHODONTIDAE

ORDER PANDERODONTIDA

Incertae Sedis:

Teridontus

FAMILY PANDERODONTIDAE

Panderodus.

ORDER PRIONIODONTIDA

UNASSIGNED GENUS

Rossodus

FAMILY MULTIOISTODONTIDAE

FAMILY OISTODONTIDAE

FAMILY PERIODONTIDAE

Periodon.

FAMILY RHIPIDOGNATHIDAE

Trincodus, Appalachignathus, Bergstroemognathus, Rhipidognathus.

FAMILY PRIONIODONTIDAE

Prioniodus, Reutterodus, Striptognathus.

FAMILY BALOGNATHIDAE (?)

FAMILY POLYPLACOGNATHIDAE

FAMILY DISTOMODONTIDAE

Distomodus.

FAMILY ICRIODELLIDAE

Icriodella.

FAMILY ICRIODONTIDAE

Icriodus.

FAMILY CYRTONIODONTIDAE

ORDER PRIONIODINIDA

UNASSIGNED GENERA

Hibbardella, Idioprionodus, Kladognathus, Phragmodus.

FAMILY CHIROGNATIDAE

Chirognathus.

FAMILY PRIONIODINIDAE

Promissum.

FAMILY BACTROGNATHIDAE

FAMILY ELLISONIIDAE

FAMILY GONDOLELLIDAE

Gondolella, Neogondolella.

FAMILY PLECTODINIDAE (paraphyletic)

Aphelognathus, Plectodina, Baltoniodus.

ORDER OZARKODINIDA

UNASSIGNED GENERA

Cordylodus

FAMILY SPATHOGNATHODONTIDAE

Ancyrodella, Ancyrognathus, Ozarkodina.

FAMILY PTEROSPATHODONTIDAE

FAMILY KOCKELELLIDAE

FAMILY POLYGNATHIDAE

Polygnathus.

FAMILY PALMATOLEPIDAE

Plamatolepis.

FAMILY ANCHIGNATHODONTIDAE

FAMILY ELECTOGNATHIDAE

FAMILY GNATHODONTIDAE

FAMILY IDIOGNATHODONTIDAE

FAMILY MESOGNATHIDAE

FAMILY CAVUSGNATHIDAE

FAMILY SWEETOGNATHIDAE