SYSTEMATIC BIOLOGY					THE VIRIDIPLANTAE
HOME	SYLLABUS	WEEKLY ASSIGNMENTS	J. SYSTEMATIC BIOLOGY	TAXA OF LIFE
THE NONVASCULAR EMBRYOPHYTES

These are nonvascular archegoniate plants (embryophytes) that seem to be related; however, as illustrated by Tudge (2000), they do not form a natural group.  The liverworts and hornworts are sister groups, but the mosses are sisters to the vascular plants. Besides the absence of vascular tissue (though some hepatophytes and mosses do have conductive tissue), they all have a dominant gametophyte stage and a more reduced or ephemeral sporophyte.  In the hepatophytes and mosses the sporophyte is determinate and parasitic on the gametophyte.  

Kenrick and Crane (1997) report that membrane-bound tetrads of spores, signature structures of embryophytes, have been found in the mid Ordovician Period (see the Geological Time Scale).  These likely were liverwort-like plants and as such did not leave any recognizable megafossils.  Early Silurian fossils of individual spores are interpreted as the appearance of mosses and hornworts, organisms that have adaptations for shedding spores over extended periods.  The transition from this "Eoembryophytic" (Kenrick and Crane 1997) flora to the vascular land plants occurred by the mid Silurian when unmistakable megafossils of plants bearing xylem appeared.

The separation of the these organisms into 3 phyla (=divisions) is drawn from Bold et al. (1987) and a modification of Schofield (1985) and Margulis and Schwartz (1998).    Indeed, the whole taxonomic scheme is drawn from those source with the modification of removing Tetraphidales, Polytrichales, and Buxbaumiales from the Bryopsida and placing them in their own respective classes within the Bryophyta. The genera given below represent examples of their orders not all-inclusive lists of genera within those orders.

Phylum Hepatophyta

CLASS MARCHANTIOPSIDA

  SUBCLASS MARCHANTIIDAE

ORDER MARCHANTIALES

Marchantia, Lunularia, Dumortiera, Conocephalum, Reboulia, Asterella, Cryptomitrium, Plagiochasma, Wiesnerella, Bucegia, Dumortiera, Neohodgsonia, Peltolepis, Monoselenium, Athalamia, Sauteria, Aitchisoniella, Exormotheca, Stephensoniella, Cyathodium, Cronisia, Corsinia, Monocarpus, Targionia.

ORDER MONOCLEALES

Monoclea.

ORDER RICCIALES

Oxymitra, Riccia, Ricciocarpos.

  SUBCLASS SPHAEROCARPIDAE

ORDER SPHAEROCARPALES

Sphaerocarpos, Geothallus, Riella.

CLASS JUNGERMANNIOPSIDA

  SUBCLASS METZGERIIDAE

ORDER HAPLOMITRIALES

Haplomitrium.

ORDER BLASIALES

Blaisia, Cavicularia.

ORDER TREUBIALES

Treubia, Apotreubia, Phyllothalia.

ORDER FOSSOMBRONIALES

Austrofossombronia, Fossombronia, Petalophyllum, Sewardiella, Allisonia, Calycularia, Noteroclada, Pellia, Sandeothallus.

ORDER METZGERIALES

Pallavicinia, Sandiothallus, Jensenia, Hattorianthus, Symphyogyna, Symphyogynopsis, Xenothallus, Moerckia, Greenothallus, Podomitrium, Makinoa, Verdoornia, Hymenophyton, 

Aneura, Cryptothallus, Riccardia, Lobatiriccardia, Mizutania, Vandiemenia, Apometzgeria, Austrometzgeria, Steereella, Metzgeria.

  SUBCLASS JUNGERMANNIIDAE

ORDER LEPICOLIALES

Ptilidium, Mastigophora, Dendromastigophora, Chaetophyllopsis, Herzogianthus.

Vetaforma, Lepicolea.

Gottschea, Pachyschistochila, Paraschistochila, Pleurocladopsis, Schistochila, Perssoniella, Eotricholea, Trichocolea, Gackstroemia, Lepidogyna, Lepidolaena, Trichocoleopsis, Neotrichocolea, Jubulopsis.

ORDER JUNGERMANNIALES

Herbertus, Triandrophyllum

Archaeophylla, Blepharostoma, Chaetocolea, Herzogiaria, Isophyllaria, Pseudolepicolea, Lophochaete, Temnoma, Trichotemnoma, Grollea.

Anisostachis, Austroscyphus, Balantiopsis, Eoisotachis, Hypoisotachis, Isotachis, Neesioscyphus, Ruzanthus.

Chiloscyphus, Clasmatocolea, Conoscyphys, Evansianthus, Geocalyx, Harpanthus, Hepatostolonophora, Heteroscyphus, Lamellocolea, Leptophyllopsis, Leptoscyphopsis, Leptoscyphus, Lophocolea, Pachyglossa, Pedinophyllopsis, Perdusenia, Pigafettoa, Platycaulis, Pseudolophocolea, Saccogyna, Saccogynidium, Stolonivector, Tetracymbaliella, Xenocephalozia, Gyrothyra, Acrochila, Pedinophyllum, Plagiochila, Plagiochilidium, Plagiochilion, Steereochila, Szweykowskia, Xenochila, Acrobolus, Austrolophozia, Enigmella, Geobelobryum, Lethocolea, Marsupidium, Tylimanthus, Arnella, Gongylanthus, Southbya.

Phycolepidozia, Calypogeia, Metacalypogeia, Acromastigum, Arachniopsis, Austrolembidium, Bazzania, Chloranthelia, Dendrobazzania, Drucella, Hyalolepidozia, Hygrolembidium, Isolembidium, Kurzia, Lembidium, Lepidozia, Megalembidium, Micropterygium, Mytilopsis, Neogrollea, Odontoseries, Paracromastigum, Protocephalozia, Pseudocephalozia, Psiloclada, Pteropsiella, Telaranea, Zoopsidella, Zoopsis.

Alobiella, Alobiellopsis, Cephalozia, Cladopodiella, Haesselia, Hygrobiella, Iwatsukia, Metahygrobiella, Nowellia, Odontoschisma, Pleurocladula, Schiffneria, Schofieldia, Trabacellula, Allisoniella E. A. Hodgs., Amphicephalozia, Cephalojonesia, Cephaloziella, Cephaloziopsis, Cylindrocolea, Kymatocalyx, Stenorrhipis, Jackiella, Adelanthus, Wettsteinia.

Anthelia.

Brevianthus, Chonecolea.

Anastrepta, Anastrophyllum, Andrewsianthus, Anomocaulis, Barbilophozia, Bragginsella, Cephalolobus, Chandonanthus, Cryptochila, Cryptocolea, Cryptocoleopsis, Denotarisia, Diplocolea, Gerhildiella, Gottschelia, Gymnocolea, Gymnocoleopsis, Hattoria, Horikwaella, Jamesoniella, Jungermannia, Lophonardia, Lophozia, Mylia, Nardia, Nothostrepta, Notoscyphus, Pisanoa, Protosyzygiella, Pseudocephaloziella, Rhodoplagiochila, Roivainenia, Scaphophyllum, Sphenolobopsis, Syzygiella, Tetralophozia, Tritomaria, Vanaea, Mesoptychia, Acrolophozia, Eremonotus, Gymnomitrion, Herzogobryum, Marsupella, Poeltia, Prasanthus, Stephaniella, Stephaniellidium, Blepharidophyllum, Diplophyllum,  Douinia, Krunodiplophyllum, Macrodiplophyllum, Scapania, Scapaniella.

ORDER PORELLALES 

Ascidiota, Macvicaria, Porella.

 

Geobeliella, Frullania, Jubula, Amphijubula, Neohattoria, Schusterella, Steerea, Bryopteris, Acanthocoleus, Acantholejeunea, Acrolejeunea, Amphilejeunea,  Anoplolejeunea, Aphanolejeunea,  Aphanotropis, Archilejeunea, Aureolejeunea, Austrolejeunea, Blepharolejeunea, Brachiolejeunea, Bromeliophila, Calatholejeunea, Capillolejeunea, Caudalejeunea, Cephalantholejeunea, Cephalolejeunea, Ceratolejeunea, Cheilolejeunea, Cladolejeunea, Cololejeunea, Colura, Crossotolejeunea, Cyclolejeunea, Cyrtolejeunea, Cystolejeunea, Dactylolejeunea, Dactylophorella, Dicladolejeunea, Dicranolejeunea, Diplasiolejeunea, Drepanolejeunea, Echinocolea, Echinolejeunea, Evansiolejeunea, Frullanoides, Fulfordianthus, Haplolejeunea, Harpalejeunea, Hattoriolejeunea, Kymatolejeunea, Leiolejeunea, Lejeunea, Lepidolejeunea, Leptolejeunea, Leucolejeunea, Lindigianthus, Lopholejeunea, Luteolejeunea, Macrocolura, Macrolejeunea, Marchesinia, Mastigolejeunea, Metalejeunea, Metzgeriopsis, Microlejeunea, Myriocolea, Myriocoleopsis, Nephelolejeunea, Neurolejeunea, Nipponolejeunea, Odontolejeunea, Omphalanthus, Oryzolejeunea, Otolejeunea, Phaeolejeunea, Physantholejeunea, Pictolejeunea, Pluvianthus, Potamolejeunea, Prionolejeunea, Ptychanthus, Pycnolejeunea, Rectolejeunea, Rhaphidolejeunea, Schiffneriolejeunea, Schusterolejeunea, Siphonolejeunea, Sphaerolejeunea, Spruceanthus, Stenolejeunea, Stictolejeunea, Symbiezidium, Taxilejeunea, Thysananthus, Trachylejeunea, Trocholejeunea, Tuyamaella, Tuzibeanthus, Verdoornianthus, Vitalianthus.

ORDER RADULALES

Radula.

ORDER PLEUROZIALES

Pleurozia

Phylum Anthocerotophyta   

CLASS LEIOSPOROCEROTOPSIDA

ORDER LEIOSPOROCEROTALES

Leiosporoceros

CLASS ANTHOCEROTOPSIDA

ORDER ANTHOCEROTALES

Anthoceros, Folioceros, Sphaerosporoceros.

ORDER NOTOTHYLADALES

Notothylas, Hattorioceros, Mesoceros, Phaeoceros, Dendroceros, Megaceros, Nothoceros.

Phylum Bryophyta

CLASS TAKAKIOPSIDA

ORDER TAKAKIALES

Takakia

CLASS SPHAGNOPSIDA

ORDER SPHAGNALES.

Sphagnum

ORDER AMBUCHANANIALES

Ambuchananiia

CLASS ANDREAEOPSIDA

ORDER ANDREAEALES.

Andreaea, Acroschima, Neuroloma.

CLASS ANDREAEOBRYOPSIDA

ORDER: ANDREAEOBRYALES  

Andreaeobryum

CLASS TETRAPHIDOSPIDA

ORDER TETRAPHIDALES.

Tetraphis, Tetrodontium.

CLASS BUXBAUMIOPSIDA

ORDER BUXBAUMIALES.

Buxbaumia, Oedipodium.

CLASS POLYTRICHOPSIDA

ORDER POLYTRICHALES.

Polytrichum, Alophosia, Atrichum, Pogonatum, Bartramiopsis, Dawsonia, Dendroligotrichum, Herbantia, Itatiella, Lyellia, Meiotichum, Notoligotrichum, Oligotrichum, Plagioraclopus, Polytrichadelphus, Polytrichastrum, Pseudatrichum, Pseudoracelopus, Psilopilum, Racelopodopsis, Racelopus, Steereobryon.

CLASS BRYOPSIDA

  SUBCLASS DIPHYSCIIDAE

ORDER: DIPHYSCIALES

Diphyscium, Muscofloeschuetzia, Theriotia.

  SUBCLASS FUNARIIDAE

ORDER: TIMMIALES

Timmia

ORDER: ENCALYPTALES

Bryobrittonia, Encalypta

ORDER FUNARIALES

Funaria, Aphanorrhegma, Brachymeniopsis, Bryobeckettia, Corynotheca,  Ochyra, Cygnicollum,  Enthosthodon, Funaria, Funariella, Physcomitrellopsis, Physcomitrium, Pyramidula, Discelium, Chamaebryum, Costesia, Gigaspermum, Lorentziella, Neoshapiella, Oedipodiella.

  SUBCLASS DICRANIDAE

ORDER GRIMMIALES

Grimmia, Neckera, Forrestroemia.

Aligrimmia, Coscindon, Coscinodontella, Dryptodon, Indusiella, Jaffueliobryum, Leucoperichaetium, Racomitrium, Schistidium, Campylostelium, Gyphomitrium, Ptychomitriopsis, Ptychomitrium, Scouleria, Tridontium, Drummondia.

ORDER ARCHIDIALES

Archidium.

ORDER:SELIGERIALES

Blindia, Brachydontium, Hymenolomopsis, Seligeria, Trochobryum, Wardia.

ORDER FISSIDENTALES

Bryoxiphium, Fissidens.

ORDER DICRANALES

Anisothecium, Aongstroemia, Aongstoempsis, Arcota, Atractylocarpus, Braunfelsia, Brothera, Bryohumbertia, Bryotestua, Camptodontium, Campylopodiella, Campylopodium, Campylopus, Chorisodontium, Cnestrum, Cryptodicranum, Dicranell, Dicranodontium, Dicranoloma, Dicranum, Eucamptodontopsis, Holomitriopsis, Holomitrium, Hygodicranum, KiaeriaI, Kingiobryum, Leptotrichella, Leucoloma, Macrodictyum, Mesotus, Microcampylopus, Mitrobryum, Muscoherzogia, Orthodicranum, Paraleucobryum, Parisia, Pilopogon, Platyneuron, Pocsiella, Polymerodon, Pseudephemerum, Pseudochorisodontium, Schiliephackea, Sclerodontium, Sphaerothecium, Steyermarkiella, Symblepharis, Werneriobryum, Bruchia, Cladophascum, Pringleella, Trematodon, Dicnemon, Eucamptodon, Cladopodanthus, Leucobryum, Ochrobryum, Schistomitrium, 

Astomiopsis, Austrophilibertiella, Bryomanginia, Cerotodon, Cheilothela, Chrysoblastella, Cladastomum, Cleistocarpidium, Crumuscus, Cygniella, Distichium, Dittrichopsis, Ditrichum, Eccremidium, Garckea, Kleioweisiopsis, Pleuriditrichum, Pleuridium, Rhamphidium, Saelania, Skottsbergia, Strombulidens, Trichodon, Tristichium, Wilsoniella, Amphidium, Cynodontium, Dichodontium, Dicraoweisia, Holodontium, Oncophorus, Oreas, Oreoweisia, Pseudohyophila, Rhabdoweisia, Verrucidens, Hypnodontopsis, Jonesiobryum, Rhachitheciopsis, Rhachithecium, Tisserantiella, Uleastrum, Zanderia.

Aulacopilum, Bryowijkia, Erpodium, Solmsiella, Venturiella, Wildia, Schistostega, Eustichia, Viridivellus.

ORDER POTTIALES

Acaulon, Aloinella, Aloinia, Anoectangium, Aschisma, Barbula, Bellibarbula, Bryoceuthospora, Bryoerythrophyllum, Calymperastrum, Calyptopogon, Chenia, Chinoloma, Crossidium, Crumia, Dialytrichia, Didymodon, Dolotortula, Erythrophyllastrum, Erythrophyllopsis, Eucladium, Ganguleea, Gertrudiella, Globulinella, Goniomitrium, Gymnostomiella, Gymnostomum, Gyroweisia, Hannediella, Hilpertia, Hymenostyliella, Hymenostylium, Hyophila, Hyophiladelphus, Hypodontium, Leptobarbula, Leptodontiella, Leptodontium, Luisierella, Microbryum, Microcrossidium, Mironia, Molendoa, Neophoenix, Pachyneuropsis, Phascopsis, Plaubelia, Pleurochaete, Pottiopsis, Pseudocrossidium, Pseudosymblepharis, Pterygoneurum, Quaesticula, Reimersia, Rhexophyllum, Sagenotortula, Saitobryum, Sarconeuron, Scopelophila, Splachnobryum, Steonia, Stonea, Streptocalypta, Streptopogon, Streptotrichum, Syntrichia, Teniolophora, Tetracoscindon, Tetrapterum, Timmiella, Tortella, Tortula, Trachycapidium, Trachydontium, Trichostomum, Triquetrella, Tuerckheimia, Uleobryum, Weisiopsis, Weissia, Weissiodicranum, Willia, Ephemerum, Micromitrium, Nanomitropsis, Seportortella, Arthrocormus, Calympres, Exostratum, Leucophanes, Mitthyridium, Octoblepharum, Syrrhopodon, Cinclidotus, Bryobartramia.

  SUBCLASS BRYIDAE

ORDER: SPLACHNALES

Aplodon, Brachymitrion, Moseniella, Splachnum, Tayloria, Tetraplodon, Voitia, Amblyodon, Leptobryum, Meesia, Neomeesia, Paludella, Catoscopium  

ORDER ORTHOTRICHALES

Bryomaltea,  Cardotiella, Ceuthotheca, Codonoblepharon, Desmotheca, Florschuetziella, Groutiella, Leiomitrium, Leptodontiopsis, Macrocoma, Macromitrium, Matteria, Mulleriella, Orthotrichum, Pentastichella, Pleurorthotrichum, Schlotheimia, Sehnemobryum, Steneobryum, Ulota, Zygodon.

ORDER:HEDWIGIALES

Braunia, Hedwigia, Hedgwigidium, Pseudobraunia, Helicophyllum, Parahacocarpus, Rhacocarpus.

ORDER BRYALES = EUBRYALES

Aulocomnium, Anacolia, Bartramia, Breutelia, Conostomum, Fleischerobryum, Flowersia, Leiomela, Philontis, Plagiopus, Quathlamba, Orthodontiym, Orthodontopsis, Acidodontium, Anomobryum, Brachymenium, Bryum, Mielichhobryum, Mniobryoides, Personia, Plagiobryum, Pleurophascum, Rhodobryum, Roellia, Rosulabryum,  Mniomalia, Phyllodrepanium,  Pseudoditrichum  Cinclidium, Cyrtomnium, Epipterygium, Leucolepsis, Mielichhoferia, Mnium, Otthomnium, Plagiomnium, Pohlia, Pseudobryum, Pseudopohlia,  Rhizomnium, Schizymenium, Synthetodontium, Trachycystis, Leptostomum.  

ORDER: RHIZOGONIALES

Hypnodendron, Cryptopodium, Goniobryum, Hymenodon, Hymenodontopsis, Leptotheca, Mesochaete, Pyrrhobryum, Rhizogonium, Calomnion, Mittenia, Bescherellia, Cyropodendron, Cyrtopus,  Franciella, Spiridens, Pterobryella, Powellia, Racopilum.

ORDER: HOOKERIALES

Achrophyllum, Cyathophorella, Cyathophorum, Dendrocyathophorum, Hookeria, Schimperobryum, Leucomium, Philophyllum, Rhynchostegiopsis, Calyptrochaeta, Crosbya, Daltonia, Distichophyllidium, Distichophyllum, Ephemeropsis, Leskeodon, Leskodontopsis, Metadistichophyllum, Adelothecium, Bryobrothera , Actinodontium, Amblytropis, Beeveria, Brymela, Callicostella, Callicostellopsis, Crossomitrium, Cyclodictyon, Diploneuron, Helicoblepharum, Hermiragis, Hookeriopsis, Hypnella, Lepidopilidium, Lepidopilum, Pilotrichidium, Pilotrichum, Sauloma, Stenodesmus, Stenodictyon, Tetrastichium, Thamniopsis, Trachyxiphium, Vesiculariopsis,  Canalohypopterygium, Catharomnion, Hypopterygium, Lopidium.

Cladomnion, Cladomniopsis, Glyphothecium, Hampeella, Ptychomnium, Tetraphidopsis, Endotrichellopsis, Euptychium, Garovaglia.

ORDER HYPNALES

Neorutenbergia, Pseudocryphea, Rutenbergia, Braithwaitea, Trachyloma

Amblystegium, Donrichardsia, Gradsteinia, Hygroamblystegium, Hypnobartlettia, Koponenia, Leptodictyum, Limbella, Ochyraea, Orthotheciella, Platylomella, Scriaromiella, Sciaromiopsis, Sinocalliergon, Vittia Cratoneuron, Sasaokaea,  Actinotuidium, Bryochena, Helodium,, Palustriella, Hylocomiastrum, Hylocomium, Leptocladiella, Leptothymenium, Leskeobryum, Macrothamnium, Meteoriella, Neodolichomitra, Orontobryum, Pleurozium,  Rhytidiadelpus, Rhytidiopsis, Schofieldiella, Rhytidium

Claopodium, Dolichomitriopsis, Fabronidium, Haplocladium, Hylocomiopsis, Leptocladium, Leptopterigynandrum, Lescuraea, Leaskeadelphus,  Leskeella, Lindbergia, Mamillariella, Miyabea, Okamuraea, Orthoamblystegium, Pseudoleskea, Pseudoleskeella, Pseudoleskeopsis, Ptychodium, Rigodiadelphus, Schwetschkea, Regmatodon.

Hordon, Heterocladium, Iwatusukiella, Myurella, Pterigynandrum, Trachyphyllum, Rigodium Abietinella, Boulaya, Cyrtohypnum, Fauriella, Pelekium, Rauiella, Thuidiopsis, Thuidium.

  Anacamptodon, Callialaria, Calliergon, Campyliadelphus, Campylium, Conardia Cratoneuropsis, Drepanocladus, Hamatocaulis, Hygrohypnum, Loeskypnum, Pictus, Pseudocalliergon, Sqnionia, Scorpidium, Straminergon, Tomentypnum, Warnstorfia.  

Aerobryum, Aerolindigia, Besia, Brachythecium, Bryhnia, Bryoandersonia, Bryostereimannia, Cirriphyllum, Clasmatodon, Cratoneurella, Eriodon, Eurhychiella, Eurhynchium, Flabellidium, Homalotheciella, Homalothecium, Isothecium, Juratzkaeella, Lindigia, Mandoniella, Meteoridium, Myuroclada, Nobregaea, Palamocladium, Plasteurhynchium, Platypnidium, Pseudopleuropus, Pseudoscleropodium, Puiggariopsis, Rhynchostegiella, Rhynchostegium, Rozea, Schimperella, Scleropodiopsis, Scleropodium, Scorpiurium, Squamidium, Steerecleus, Stenocarpidiopsis, Trachybryum, Zelometeorium.  

Catogoniopsis, Entodontopsis, Eulacophyllum, Juratzekaea, Pilosium, Sciuroleska, Stenocaridium, Stereophyllum. Austinia, Helicodontium, Macrgregorella, Merrilliobryum, Myrinia, Nematocladia, Dimerodontium, Fabronia, Ischyrodon, Levierella, Rhizofabronia.

Aerobryidium Aerobryopsis, Ancistrodes Barbella, Barbellopsis, Chrysocladium, Cryphaeophilum, Cryptopapillaria, Diaphanodon, Duthiella, Floribundaria, Fleisch, Lepyrodontopsis, Meteoriopsis, Meteorium, Neodicladiella, Neonoguchia, Pseudospiridentopsis, Pseudotrachypus, Sinskea, Toloxis, Trachycladiella, Trachypodopsis, Trachypus, Plagiothecium. Brachelyma, Dichelyma, Fontinalis,  Entodon, Erythrodontium, Mesondon, Pylaisiobryum, Climacium, Pleuroziopsis.

  Andoa, Bardunovia, Breidleria, Bryocrumia, Bryosedwickia, Callicladium, Calliergonella, Campylophyllum, Caribaehypnum, Chrysohypnum, Crepidophyllum, Ctenidiadalphus, Caribaeohypnum, Chrysohypnum, Crepidophyllum, Ectropotheciella, Ectropotheciopsis, Ectropothecium, Elharveya, Giraldiella, Gollania, Herzogiella, Homomallium,  Hondaella, Horridohypnum, Hycomium, Hypnum , Irendia, Isopterygiopsis, Isopterygium, Leiodontium, Macrothamniella, Mahua, Microcteidium, Mittenothamnium, Nanothecium, Orthothecium, Phyllodon, Plagiotheciopsis, Platydictya, Platygyriella,  Platygyrium, Podperaea, Pseudotaxiphyllum, Ptilium,  Pylaisiella, Rhacopilopsis, Rhizohypnella, Sclerohypnum, Serpoleskea, Stenotheciopsis, Stereodontopsis, Syringothecium, Taxiphyllopsis, Taxiphyllum, Trachythecium, Tripterocladium, Vesicularia, Wijkiella.  

Catagonium, Chaetomitriopsis, Chaetomitrium, Dimorphocladon, Symphyodon, Acanthorrhynchium, Acritodon, Acropotrium, Allionrllopsis, Aptychella, Aptychopsis, Brotherella, Chinostomum, Clastobryella, Clastrobryophilum, Clastobryopsis, Clastobryum, Colobodontium, Donnellia, Foreauella, Gammiella, Hagniella, Heterophyllium, Isocladiella, Leptoischyrodon, Macrohymenium, Mastopoma, Meiotheciella, Papillidiopsis, Paranpiacabaea, Potamium, Pseudohypnella, Pterogonidium, Pterogoniopsis, Pylaisiadelpha, Pylaisiopsis, Radulina, Rhaphidostichum, Scaderella, Schroeterella, Sematophyllum, Struckia, Taxitheliella, Taxithellia, Taxithelium, Timotimius, Trichosteleum, Trismegistia, Trolliella , Warburgiella, Wijkia.  

Hydropogon, Hydropogonella, Eumyrurium, Myrurium, Oedicladium, Palisadula, Piloecium Cryphaea,, Cryphidium, Cyptodon, Cyptodontopsis, Dendroalsia, Dendrocryphaea, Dendropogonella, Pilotrichopsis, Schoenobryum, Sphaerotheciella, Prionodon, Antitrichia, Dozya, Eoleucodon, Felipponea, LeucodonSchägr, Pterogonium, Scabridens, Calyptothecium, Cryptogonium, Henicodium, Hildebrandtiella, Horikawea, Jaerernia, Micralsopsis, Muellerobryum Neolindbergia, Orthorrhynchidium, Orthostichidium, Orthostichopsis, Ostewaldiella, Penzigiella, Pireella, Psuedopterobryum, Pterobryidium, Pterobryon, Pterobryopsis, Pulchrinodus, Renauldia, Rhabdodontium, Spiridentopsis, Smphysodon, Symphysodontella, Phyllogonium, Orthorrhynchium, Dichelodontium, Lepyrodon.  

Baldwiniella, Bissetia, Btyolawtonia, Caduciella, Crassiphyllm, Cryptolepton, Curvicladium, Dixonia, Dolichomitra, Handeliobryum, Himantocladium, Homalia, Homalidelphus, Homalidendron, Hydrocryphea, Isodrepanium, Metaneckera, Neckera, Neckeropsis, Meomacounia, Noguchiodendron, Pendulothecium, Pinnatella, Porotrichopsis, Porotrichum, Thamnobryum, Touwia. Alsia, Forsstroemia, Leptodon, Taiwanobryum,  Acrocladium, Camptochaete, Fifea, Lembophyllum, Neobarbella, Orthostichella, Pilotrichella, Weymouthia,  Anomodon, Bryonorrisia, Chileobryon, Curviramea, Haplohymenium, Herpetineuron, Schwetschkeopsis,  Thelia, Microtheciella, Sorapilla.

This page is maintained by Jack R. Holt & Carlos A. Iudica.  Last modified: 03/17/08 .